Search bioRxiv⌕ Search

Biology subjects

Ceravolo, L.

Publications and source records attributed to Ceravolo, L..

At least 19 recordsLinked to original sources

Distinct experience-dependent reorganization of rhythmic syllable tracking in 6-month-old term and preterm infants

Preterm birth is associated with an increased risk of later language difficulties, yet whether and how prematurity alters the brain's ability to track the rhythmic structure of speech has remained understudied. Using EEG, we tracked neural entrainment to delta- and theta-rate syllable sequences produced by the infant's mother or an unfamiliar speaker in term and preterm 6-month-old infants, modeling how oscillatory responses evolved over the course of a testing session. Term infants progressively strengthened frequency-matched delta tuning over bilateral frontal regions; preterm infants did not. This divergence took two distinct forms: a change in frequency selectivity over left frontal channels, where term infants sharpened and preterm infants blurred, and a shift in band weighting in other regions. The frontal delta difference emerged only for the unfamiliar speaker. Preterm birth thus alters how frequency-specific tuning is refined during exposure, identifying a candidate neural pathway with implications for early language learning.

neuroscience↗

Delta rhythm and voice familiarity organize neural entrainment in the infant brain

The delta frequency band (~0.5-3.5 Hz) represents a phylogenetically conserved tempo of biological communication, and neural entrainment to this timescale would be a key mechanism through which the brain might generate temporal predictions. While entrainment at different levels has been characterized in adults, the mechanisms of auditory temporal prediction remain poorly studied across developmental trajectories, particularly regarding the roles of stimulus rhythmicity, frequency specificity, and voice familiarity. Using high-density EEG, we examined neural entrainment in thirty 6-month-old infants exposed to vocally rhythmic syllables produced by their mother or a stranger at delta (2 Hz) or theta (4 Hz) rates, as well as a non-rhythmic control. Auditory temporal regularity elicited a broad frontotemporal entrainment response. Critically, delta-band entrainment was frequency-specific across frontal and temporal clusters, with right-frontal selectivity progressively strengthening throughout the entrainment period as expected if this region actively refines a temporal model of the input rather than passively resonating to it; no significant theta-band specificity was observed. Entrainment was further shaped by voice familiarity: the left temporal cortex entrained selectively to the mother's voice, whereas the right frontal cluster responded selectively to the stranger's voice. This is consistent with an implementation in which familiarity-based processing within mid-temporal language-related regions interfaces with novelty-sensitive frontal mechanisms, and reflects the ongoing updating of internal predictive models rather than passive sensory registration. The co-occurrence of frequency-specific delta entrainment and experience-dependent hemispheric asymmetries reveals a neural functional network for hierarchical temporal predictions, shaped by early auditory experience and already in place at 6 months.

neuroscience↗

The integration of prosody and semantics in non-literal speech: A voxel-wise encoding model approach using large language models

Irony and sarcasm are complex forms of non-literal language that hinge on a misalignment between surface meaning and speaker intent, requiring listeners to integrate contextual, semantic, and prosodic cues. While prior neuroimaging studies have implicated a broad network--including the temporal cortex, the inferior frontal gyrus, and the medial prefrontal cortex--in the comprehension of ironic and sarcastic speech, the precise neural mechanisms underlying the integration of semantic and prosodic information remain unclear. In the present study, we addressed this gap by employing voxel-wise encoding models to systematically identify brain regions specifically involved in combining prosodic and semantic cues during non-literal language comprehension. Participants listened to naturalistic auditory dialogues in which both discourse context and target utterance semantics and prosody were systematically manipulated. We derived custom text embeddings using transformer-based models to capture context-sensitive semantic representations of ironic statements, alongside acoustic features characterizing affective prosody. Ridge regression models were fitted to predict BOLD responses at the voxel level using semantic, prosodic, and combined features, and we identified integration as voxels in which each modality contributed predictive information beyond the other, using a permutation-based conjunction test. The regions integrating prosody and semantics depended on whether discourse context was modeled: integration was confined to the bilateral temporal speech cortex when statements were encoded in isolation, but additionally engaged the left inferior frontal gyrus pars orbitalis (IFGorb) when each statement was weighted by its relevance to the preceding context. These findings indicate that the left IFGorb integrates prosody with context-dependent meaning, engaging beyond the temporal speech cortex specifically when comprehension requires combining semantic, prosodic, and contextual cues--as in irony and sarcasm.

neuroscience↗

Shared book reading promotes experience-dependent autonomic synchrony in parent-preterm infant dyads

Preterm birth is associated with alterations in early caregiver-infant regulation, with potential consequences for socio-emotional and physiological development. However, the mechanisms through which early interactional experience shapes these processes remain unclear. Here, we tested whether a structured dyadic intervention could modify co-regulatory dynamics across physiological, behavioral, and relational levels. Fifty-four 7-month-old preterm infants and their parents were assigned to either a shared book reading intervention (n = 22) or an active control condition based on a shared building activity (n = 32) and compared with 39 full-term infants. The intervention consisted of an 8-week program of shared book reading, designed to structure parent-infant interaction. Physiological synchrony was assessed at the dyadic level, alongside infants autonomic regulation and cardiovascular signal complexity. Behavioral engagement and parental attachment representations were also evaluated. Results showed that mother-infant physiological synchrony emerged selectively within the interactional context trained by the intervention and only in the intervention group. This context-specific synchrony was accompanied by modulation of vagal activity and increased cardiovascular complexity in preterm infants, consistent with enhanced flexibility of autonomic control. At the behavioral and relational levels, intervention infants showed increased initiating joint attention, while parents reported higher secure attachment. These findings support a model of experience-dependent early synchrony, in which repeated dyadic interaction through shared book reading shapes the coupling between interpersonal coordination and individual physiological regulation. By linking synchrony, autonomic flexibility, and social engagement, this study identifies a mechanism through which early caregiving experience can organize developmental trajectories following prematurity.

developmental biology↗

Neural Integration of Affective Prosodic and Semantic Cues in Non-literal Forms of Speech Understanding

Emotions in speech are conveyed through both semantics and prosody--voice melody, yet how listeners integrate these cues in the brain remains unclear. We investigated non-literal forms of speech, such as irony and sarcasm, where understanding beyond literal meaning relies on the dynamic interplay between affective prosodic and semantic cues, alongside theory of mind (ToM). We used functional magnetic resonance imaging while participants listened to short dialogues between two characters, which varied in prosody and semantics to convey either literal or non-literal meanings. Behaviorally, semantics and prosody interacted in shaping participants evaluations, although the data suggest a prosody dominance effect. Non-literal speech engaged a distributed network including the bilateral inferior frontal gyrus, temporal speech regions, and ToM areas, with ROI analyses revealing heterogeneous prosody-semantics integration profiles across regions and tasks. Taken together, our data clarify the behavioral and neural underpinnings of the integration of prosody and semantics in non-literal speech and open new venues in this field. Author noteThe authors made the following contributions. Adrien Wittmann: Conceptualization, Methodology, Data curation, Formal analysis, Software, Investigation, Validation, Visualisation, Writing - Original Draft Preparation, Writing - Review & Editing; Leonardo Ceravolo: Methodology, Software, Writing - Review & Editing; Audrey Mayr: Conceptualization, Software, Methodology, Investigation, Validation; Didier Grandjean: Funding acquisition, Supervision, Conceptualization, Writing - Review & Editing.

neuroscience↗

Attentional focus and emotion modulate voice recognition deficits in cerebellar stroke patients

The cerebellum, long regarded as a motor structure, is increasingly recognized for its role in higher-order cognitive and socio-emotional functions. Its contribution to vocal emotion decoding, however, remains insufficiently understood. While prior work has linked the cerebellum to attentional control and predictive coding, direct evidence for its role in modulating prosody recognition under explicit versus implicit attentional demands is lacking. This study investigated how cerebellar stroke--controlled for time since stroke in months, and with an equal number of left- and right-lateralized lesions--alters vocal emotion processing depending on attentional focus. We aimed to disentangle sensory encoding and integration from decisional contributions of the cerebellum by combining behavioral analyses, drift diffusion modeling (DDM), and functional MRI in cerebellar stroke. Fifteen patients with chronic, isolated cerebellar stroke and fifteen matched controls performed two tasks on vocal stimuli expressing anger, happiness, or neutrality. In the explicit task, participants categorized the expressed emotion; in the implicit task, they categorized speaker gender while ignoring emotional tone. Behavioral performance was analyzed using mixed-effects logistic regression and DDM (angle model). Functional MRI analyses included conventional contrasts as well as model-based regressors derived from behavioral and computational parameters. Patients showed lower accuracy than controls, with deficits particularly pronounced in the explicit task. Performance also varied by emotion: gender recognition for angry voices (implicit processing) was relatively preserved, whereas happy and neutral voices were more vulnerable. Contrary to predictions, DDM parameters did not differ between groups. Instead, task effects dominated: implicit processing yielded higher drift rates, larger boundary separation, faster non-decision times, and stronger attentional focus compared to explicit recognition, suggesting that explicit evaluation imposes additional cognitive costs. At the neural level, patients recruited extended networks during explicit processing, including orbitofrontal cortex, amygdala, anterior insula, and cerebellar lobule IX, alongside cerebello-cortical tracts. Implicit processing was associated with more restricted activations, particularly in frontal opercular and cerebellar regions (lobule IX). Model-based analyses further revealed that successful categorization in patients relied on the left inferior frontal gyrus, inferior parietal lobule, and pre-supplementary motor area, although these effects were not significant when controlling for time since stroke. Our findings demonstrate that the cerebellum does not primarily shape decision dynamics but optimizes sensory representations of prosodic cues for downstream evaluation. When predictive tuning is compromised, patients maintain intact decision processes yet rely on compensatory cortical recruitment, particularly during explicit tasks. This pattern supports predictive coding accounts of cerebellar function in socio-emotional communication and highlights the need to consider subtle socio-affective deficits in cerebellar patients.

neuroscience↗

Sensitivity of the human temporal voice areas to nonhuman primate vocalizations

In recent years, research on voice processing in the human brain, particularly the study of temporal voice areas (TVA), was dedicated almost exclusively to conspecific vocalizations. To characterize commonalities and differences regarding primate vocalization representations in the human brain, the inclusion of closely related nonhuman primates, namely chimpanzees and bonobos, is needed. We hypothesized that neural commonalities would depend on both phylogenetic and acoustic proximities, with chimpanzees ranking closest to Homo. Presenting human participants (N=23) with the vocalizations of four primate species (rhesus macaques, chimpanzees, bonobos and humans) and regressing-out relevant acoustic parameters using three distinct analyses, we observed within-TVA, sample-specific, bilateral anterior superior temporal gyrus activity for chimpanzee vocalizations compared to: all other species; nonhuman primates; human vocalizations. Within-TVA activity was also observed for macaque vocalizations. Our results provide evidence for subregions of the TVA that respond principally, but not exclusively, to phylogenetically and acoustically close nonhuman primate vocalizations, namely those of chimpanzees.

neuroscience↗

Neural dynamics of induced vocal tract vibrations during vocal emotion recognition

Despite a large corpus of literature in psychological and brain mechanisms on emotional prosody perception, the perspective of embodied cognition in these mechanisms have been largely neglected. Here we investigated the influence of induced bodily vibrations on the categorization of ambiguous emotional vocalizations using event-related potentials (ERPs). Emotional voices were morphed between a fearful expression with the speakers identity-matching angry expression, creating blends of emotions in each voice. Emotional congruent and incongruent vibrations were delivered on the skin close to the vocal cords. Congruent with our hypotheses, behavioural results revealed that induced vibrations skewed the participants emotional ratings by biasing responses towards the vibrations emotion. ERPs indicated that N100 and P200 components subtending the early processing of emotional prosody were significantly modulated by induced vibrations in the congruent setting, considered as a facilitation effect for emotion recognition at early stages of processing. A modulation of the late positive component was also observed in the incongruent setting, suggesting an error processing mechanism. Source reconstruction highlighted effects of vibration types in prefrontal, motor, somatosensory, and insular cortices. Our results suggest that voice-associated vibrations may play a significant role in vocal emotion processing and recognition through an embodied mechanism.

neuroscience↗

Neural correlates of approach and avoidance tendencies toward physical activity and sedentary stimuli: An fMRI study

Automatic tendencies toward physical activity and sedentary stimuli are involved in the regulation of physical activity behavior. However, the brain regions underlying these automatic tendencies remain largely unknown. Here, we used an approach-avoidance task and magnetic resonance imaging (MRI) in 42 healthy young adults to investigate whether cortical and subcortical brain regions underpinning reward processing and executive function are associated with these tendencies. At the behavioral level, results showed more errors in avoidance behavior following sedentary stimuli than physical activity stimuli. At the brain level, avoidance behavior following sedentary stimuli was associated with more activation of the motor control network (dorsolateral-prefrontal cortex, primary and secondary motor cortices, somatosensory cortex). In addition, increased activation of the bilateral parahippocampal gyrus -- and structural deformation of the right hippocampus - were associated with a tendency toward approaching sedentary stimuli. Together, these results suggest that avoiding sedentary stimuli requires higher levels of behavioral control than avoiding physical activity stimuli.

neuroscience↗

Differential Engagement of Associative-Limbic and Sensorimotor Regions of the Cerebellum and Basal Ganglia in Explicit vs. Implicit Emotional Processing

Emotional prosody processing involves a widespread network of brain regions, but the specific roles of the cerebellum and basal ganglia in explicit and implicit tasks are not well known or understood. This study investigated how the cerebellum and basal ganglia contribute to explicit (emotion categorization) and implicit (gender categorization) processing of emotional prosody, namely when attention is directly versus implicitly oriented towards the emotion of the voice stimuli, respectively. Twenty-eight healthy French-speaking participants (average age: 65 years old) underwent high-resolution functional MRI while performing explicit and implicit vocal emotion processing tasks. Neuroimaging results revealed--and replicated--that both tasks recruited a widespread network, including the superior temporal cortex, inferior frontal cortex, primary motor and somatosensory cortices, basal ganglia, and cerebellum. The explicit task elicited stronger activations in the basal ganglia (caudate nucleus, putamen) and cerebellar regions (Crus I/II, lobules VI, VIIb, and X), consistent with higher cognitive control demands. In contrast, the implicit task was associated with activations in cerebellar lobules IV-V, VI, VIII, and IX, along with the thalamus. Regression-based functional connectivity analyses further demonstrated stronger connectivity between the right cerebellar lobule IX and the putamen, as well as the cerebellar vermis (XII), particularly during implicit processing. These findings highlight the distinct contributions of the cerebellum and basal ganglia to emotional prosody processing, with explicit tasks engaging associative and cognitive control networks, while implicit tasks rely more on sensorimotor and automatic neural processing mechanisms.

neuroscience↗

Functional and causal neural mechanisms of human voice perception in noisy situations

Human communication entails an efficient way of simultaneously processing voice and reducing the impact of environmental noise. By manipulating background noise, we aimed at clarifying the neural mechanisms allowing voice comprehension in noisy situations. Our results point to spatial and temporal coexistence of lateral and medial temporal cortex networks when voice is easily detected in highly noisy conditions, revealing the necessary neural underpinnings of human communication in realistic situations.

neuroscience↗

Neural correlates of embodied and vibratory mechanisms associated with vocal emotion production

Despite a large body of literature on the psychological and brain mechanisms of vocal emotion perception, less is known on expression and production mechanisms, especially the vibrations originating in the vocal cords and their role in emotional voice production. In the present study, we aimed to fill this gap. Participants had to produce angry, happy and neutral emotional vocalizations in different production conditions ( normal, whisper, articulate). An accelerometer recorded the vibrations on the throat, close to the vocal folds. Results showed effects of the Emotion factor with activations in the bilateral temporal voice areas, the inferior frontal gyri as well as motor and supplementary motor areas. The Production factor and its interaction with Emotion revealed significant effects in motor, somatosensory cortices, insula and inferior frontal cortex. Exploratory analysis of the brain correlates of emotional vocal tract vibrations specific to normal voice production showed significant correlations with brain regions involved in interoceptive processing (insula, inferior frontal cortex, cerebellum). Our results highlight the crucial role of vibro-tactile body resonances in vocal emotion production that might play an important role for the interoceptive phenomena involved in the representation of our own emotions such as in emergent feelings related to emotional vocal production.

neuroscience↗

Model-based frontal cortex analysis reveals origins of human non-verbal communication

The ability to process verbal language seems unique to humans and relies not only on semantics but on other forms of communication such as affective vocalisations, that we share with other primate species--particularly great apes (Hominidae). To better understand these processes at the behavioural and brain level, we asked human participants to categorize vocalizations of four primate species including human, great apes (chimpanzee and bonobo), and monkey (rhesus macaque) during MRI acquisition. Classification was above chance level for all species but bonobo vocalizations. Imaging analyses were computed using a participant-specific, trial-by-trial fitted probability categorization value in a model-based style of data analysis. Model-based analyses revealed the implication of the bilateral orbitofrontal cortex and inferior frontal gyrus pars triangularis (IFGtri) respectively correlating and anti-correlating with the fitted probability of accurate species classification. Further conjunction analyses revealed enhanced activity in a sub-area of the left IFGtri specifically for the accurate classification of chimpanzee calls compared to human voices. Our data therefore reveal distinct frontal mechanisms that shed light on how the human brain evolved to process non-verbal language. Author contributionsCD and LC were involved in every steps of the study including experimental design, programming, data acquisition, data analysis and redaction of the first draft of the manuscript and subsequent editing. EP helped design the model-based MRI analyses and interpret the results. TG and DG were involved in the experimental design and study planification. All authors wrote and edited the manuscript.

neuroscience↗

Categorization and discrimination of human and non-human primate affective vocalizations: investigation of the frontal cortex activity through fNIRS.

Many species, including humans and non-human primates, react differently to threatening or pleasant situations. Because of its adaptiveness, recognizing affective signals is likely to be reflected in a capability of modern humans to recognize other closely related species call content. However, at both behavioural and neural levels, only few studies have used a comparative approach to understand affective decoding processes in humans, particularly with respect to affective vocalizations. Previous research in neuroscience about the recognition of human affective vocalizations has shown the critical involvement of temporal and frontal regions. In particular, frontal regions have been reported as crucial in the explicit decoding of vocal emotions especially in different task complexity such as discrimination or categorization. The aim of this study using functional Near Infrared Spectroscopy (fNIRS) was to specifically investigate the neural activity of the inferior frontal cortex pars triangularis (IFGtri) and the prefrontal cortex (PFC) underlying categorization (A versus B) and discrimination (A versus non-A) mechanisms of positive and negative affects in human, great apes (chimpanzee and bonobo), and monkey (rhesus macaque) vocalizations. We also analysed participants behavioural responses and correlated them with the recorded frontal activations. While performing the tasks, fNIRS data revealed a clear distinction between the two frontal regions, with a general positive activation of IFGtri compared to a decrease of PFC activity. We also found a modulation of IFGtri and PFC activations depending on both the species considered and on task complexity; with generally more activity in the IFGtri during discrimination compared to categorization, and a more intense decrease of the PFC in categorization compared to discrimination. Behaviourally, participants recognized almost all affective cues in all species vocalizations at above chance levels in the discrimination task (except for threatening bonobo calls). For categorization, they mostly correctly identified at levels significantly above chance affective contents in human and great ape vocalizations but not in macaque calls. Overall, these findings support the hypothesis of a pre-human origin of affective recognition processing inherited from our common ancestor with other great apes and processed in the frontal cortex. Our results also highlight behavioural differences related to task complexity, i.e. between categorization and discrimination processes, and the differential involvement of the PFC and the IFGtri, which seems necessary to explicitly decode affects in all primate vocalizations.

neuroscience↗

Disrupting inferior frontal cortex activity alters affect decoding efficiency from clear but not from ambiguous affective speech

The evaluation of socio-affective sound information is accomplished by the primate auditory cortex in collaboration with limbic and inferior frontal cortex (IFC)--often observed during affective voice classification. Partly opposing views have been proposed, with IFC either coding cognitive processing challenges in case of sensory ambiguity or representing categorical object and affect information for clear voices. Here, we presented clear and ambiguous affective speech to two groups of human participants during neuroimaging, while in one group we inhibited right IFC activity with transcranial magnetic stimulation. IFC activity inhibition led to faster affective decisions, more accurate choice probabilities, reduced auditory cortical activity and increased fronto-limbic connectivity for clear affective speech. This indicates a more intermediate functional property of the IFC than assumed--namely with normal activity representing a more deliberate form of affective sound processing (i.e., enforcing cognitive analysis) that flags categorical sound decisions with precaution (i.e., representation of categorical uncertainty). TeaserInferior frontal cortex enforces cognitive analyses during affect decisions with different levels of sensory ambiguity.

neuroscience↗

Frontoparietal, cerebellum network codes for accurate intention prediction in altered perceptual conditions

Integrating and predicting intentions and actions of others are crucial components of social interactions, but the behavioral and neural underpinnings of such mechanisms in altered perceptual conditions remain poorly understood. We demonstrated that expertise was necessary to successfully understand and evaluate communicative intent in spatially and temporally altered visual representations of music plays, recruiting frontoparietal regions and several sub-areas of the cerebellum. Functional connectivity between these brain areas revealed widespread organization, especially in the cerebellum. This network may be essential to assess communicative intent in ambiguous or complex visual scenes.

neuroscience↗

Basal ganglia and cerebellar contributions to vocal emotion processing: a high resolution fMRI study

Until recently, brain networks underlying emotional voice prosody decoding and processing were focused on modulations in primary and secondary auditory, ventral frontal and prefrontal cortices, and the amygdala. Growing interest for a specific role of the basal ganglia and cerebellum was recently brought into the spotlight. In the present study, we aimed at characterizing the role of such subcortical brain regions in vocal emotion processing, at the level of both brain activation and functional and effective connectivity, using high resolution functional magnetic resonance imaging. Variance explained by low-level acoustic parameters (fundamental frequency, voice energy) was also modelled. Wholebrain data revealed expected contributions of the temporal and frontal cortices, basal ganglia and cerebellum to vocal emotion processing, while functional connectivity analyses highlighted correlations between basal ganglia and cerebellum, especially for angry voices. Seed-to-seed and seed-to-voxel effective connectivity revealed direct connections within the basal ganglia especially between the putamen and external globus pallidus and between the subthalamic nucleus and the cerebellum. Our results speak in favour of crucial contributions of the basal ganglia, especially the putamen, external globus pallidus and subthalamic nucleus, and several cerebellar lobules and nuclei for an efficient decoding of and response to vocal emotions.

neuroscience↗

Human temporal voice areas are sensitive to chimpanzee vocalizations

Withdrawal StatementThe authors have withdrawn this manuscript due to a duplicate posting of manuscript number BIORXIV/2025/677258. Therefore, the authors do not wish this work to be cited as reference for the project. If you have any questions, please contact the corresponding author. The correct preprint can be found at doi: 10.1101/2025.09.19.677258

neuroscience↗