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Identification of the C9-Hydrogenase for 9,17-Dioxo-1,2,3,4,10,19-Hexanorandrostan-5-oic Acid (9,17-DOHNA) and the 7α-Dehydratase Essential for Initiating β-Oxidation of the B-, C-, and D-Rings in Steroid Degradation by Comamonas testosteroni TA441

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