TY - RPRT TI - Darwinian fitness, its directional derivative, and Hamilton's rule for limited dispersal with class structure under within and between generation environmental stochasticity AU - Lehmann, L. PY - 2026 DO - 10.64898/2026.05.05.722983 UR - https://www.biorxiv.org/content/10.64898/2026.05.05.722983v1 ID - 10.64898/2026.05.05.722983 ER -